1State Key Laboratory of Precision Measurement Technology and Instruments, Department of Precision Instrument, Tsinghua University, Beijing 100084, China
2School of Life Sciences, Tsinghua University, Beijing 100084, China
3Senior Department of Orthopedics, The Fourth Medical Center of Chinese PLA General Hospital, Beijing 100853, China
4IDG/McGovern Institute for Brain Research, Tsinghua University, Beijing 100084, China